The Command Center is the central operations room of NERV Headquarters. It's appearance is reminiscent of the bridge of a military ship. Seven core NERV staff members gather there, including Commander Gendo Ikari, Vice Commander Kouzou Fuyutsuki, Project E lead Dr. Ritsuko Akagi, and Operations Director Misato Katsuragi.
During combat operations, all orders are issued from this location, which also houses the original MAGI Super-Computer System. Eventually this location is breached and heavily damaged by Zeruel, the Fourteenth Angel. An auxiliary Command Center of identical construction is used following the incident.
Based on Neon Genesis Evangelion Platinum: 04 DVD Booklet